HYBRIDS INVOLVING LEOPARDS AND JAGUARS

JAGLION

A jaglion is a hybrid between male jaguar and lioness. There was a claim of a lion x black jaguar cross (male) seen in the company of an alleged tiger x black jaguar cross (female) in Maui, Hawaii. The supposed lion x black jaguar's description matches that of an African lion. It was described as having a scarred, puffy grey face, short, thick jet black mane on its head and around its neck, extending around the ears and underneath the chin. Its body was solid dark tawny and the tail had a black tuft. The identification of lion x jaguar hybrid was made on facial characteristics. Melanism is a dominant trait in jaguars and its interaction with lion genes was unknown until 2006. The grey face was probably due to poor light. Non-specialists frequently misidentify big cats and erroneously identify them as hybrids. Tigers have never been hybridised with jaguars. The two sighted animals were probably an escaped African lion and lioness.

Known jaglion hybrids have chocolate rosettes on a tawny background colour. The jaguar x lion hybrid below is a preserved individual photographed at the Natural History Museum, Tring, England. The hybrid has the lion's background colour, jaguar-like rosettes and the powerful build of the jaguar. There are no definite records of jaguar/lion hybrids prior to those born at Barrie, Ontario. Since the specimen at Tring matches the description of the lijagulep (Congolese Spotted Lion) it may be that animal. Hemmer was apparently unsure whether the museum specimen he examined was a jaguar/lion hybrid, as stated on the Tring specimen's label, or a leopard/jaguar x lion hybrid. Hemmer (1968) described a mounted specimen as a presumed lion x jaguar or lion x (jaguar x leopard) hybrid, being reddish-yellow, paler than a jaguar, with brown spots. (Photos below)

On April 9, 2006, two Jaglions were born at Bear Creek Wildlife Sanctuary, Barrie (north of Toronto), Ontario following an accidental mating. Jahzara (female,) and Tsunami (male) were the result of a mating between a black male jaguar (Diablo) and a lioness (Lola). The sanctuary does not deliberately breed animals and keepers believed Lola would not conceive. As an added precaution, the pair were separated when Lola came into oestrus. They had been hand-raised together and were inseparable, becoming anxious and depressed every time separation was attempted. Lola pined during attempted separations. Jahzara is melanistic, a dominant trait in jaguars, while Tsunami is spotted. It was not previously known how the jaguar's melanism gene would interact with lion colouration genes. Because female big cat hybrids are frequently fertile, Jahzara could theoretically be bred back to purebred lions to introduce the gene for melanism into that species.

Although displayed as a hybrid of jaguar and lion, the exhibit below is very probably the famous Congolese Spotted Lion, being a 3-way hybrid bred from a jaguar/leopard female mated to a lion.

 

JAGULEP (JAGLEOP, LEPJAG, LEGUAR)

Crosses of both male jaguar to female leopard and of male leopard to female jaguar have been reported.

In The Field No 2887, April 25th, 1908, Henry Scherren wrote, "In a paper on the breeding of the larger Felidae in captivity (P.Z.S [Proceedings of the Zoological Society]., 1861, p. 140), A.D. Bartlett stated : "I have more than once met with instances of the male jaguar (F. onca) breeding with a female leopard (F. leopardus). These hybrids were also reared recently in Wombell's well known travelling collection. I have seen some animals of this kind bred between a male black jaguar and a female Indian leopard:-the young partook strongly of the male being almost black." (It is now known that melanism in the jaguar is semi-dominant, those with one copy of the gene are almost black, while those with 2 copies are fully melanistic)

In The Field No 2887, April 25th, 1908, Henry Scherren wrote, "In Barnabos menagerie (in Spain) a jaguar threw two cubs by a black panther, one resembled the dam, but was somewhat darker, the other was black with the rosettes of the dam showing. (Zoolog. Gart., 1861, 7)" (Since melanism in the panther (leopard) is recessive, the jaguar would either have been black or be a jaguar-black leopard hybrid itself, carrying the recessive gene.) Scherren continued "The same cross, but with the sexes reveresed, was noted, by Professor Sacc (F) of Barcelona Zoo (Zool Gart, 1863, 88) "The cub a female was grey: she is said to have produced two cubs to her sire; one like a jaguar, the olther like the dam. Herr Rorig expressed his regret that the account of the last two cases mentioned lacked fulness and precision.

Hemmer (1966, 1968) described hybrids from a male jaguar and female leopard as about the size of the jaguar. The neck and legs were rather short and thick. The background colour was dark fawn, grey-brown, iron-grey or olive coloured. The rosettes on the flank were usually much attenuated. Matings of the female jaguar-leopard hybrids to a male jaguar resulted in 2 litters, each with 2 offspring. He noted that a jaguar-leopard also produced an offspring when mated to a male lion (the famous Congolese Spotted Lion hoax).

Sacc (1863) and H Scherren (1908) both reported that a grey-coloured female jaguar-leopard hybrid was mated back to her jaguar father. She produced 2 healthy cubs (jag-lepjags), one resembling the jaguar and one resembling the leopard.

C Pohle (1969) reported a jaguar-leopard hybrid litter comprising 1 female and 2 males. One male died, but at 9 months the surviving hybrids were larger than their leopard mother. At 1.5 years they were intermediate between leopard and jaguar in height at the shoulders. A further jaguar-leopard hybrid was later produced from the same parents.

H Windischbauer (1968) reported a mating of a 6 year old male African leopard (P pardus pardus) with a 2.5 year old female jaguar. Hellbrun Zoo was criticised for the hybrids offered as attractions although these were popular with visitors. Heinrich Windischbauer (director until 1976) offered visitors the "Jagupard". The mating, at Hellbrunn Zoo, Salzburg, Austria, produced 2 vigourous cubs in 1966. One cub was smaller in size and darker in colour than the other. At 6 months old, the cubs were both considerably stronger than leopards or jaguars of the same age.

The Long Island Ocelot Club (LIOC) newsletter of Jan/Feb 70 carried a piece by Mary Ellen Tracy who planned to breed jaguar x leopard hybrids. She had a male jaguar called Kwai Mao and a 2 year old Asian leopardess called Takala. She hoped to cross the jaguar and leopard to yield jaguards and test the hybrids for sterility. "We wish to test the differences between these two species to see if perhaps they are not as great as many believe."

The Long Island Ocelot Club (LIOC) newsletter of May/Jun 1976 noted "There are also many black and spotted leopards in [Robert Baudy's] collection, plus a hybrid Lepjag (a cross between a jaguar and a leopard).". In its newsletter for Sept/Oct 1977, LIOC added more information on the hybrid: For six years [Gladys Lewis] has been raising almost all the exotic cats born at Robert Baudy's Rare Feline Breeding Compound in Center Hill, Fla. One interesting aspect of her cat raising methods is the devotion her cats extend to her, even after they leave her. Gladys raised an Indian leopard (male) with a spotted jaguar female from kittenhood in her home, when they matured, not knowing they were from different continents and not the same species, they mated and eventually produced four litters of hybrids called Lep-jag. The first one is now over two years old, Suma by name, and he is still a growing boy at 150 pounds plus. His front end seems to be jaguar, (large head, wide front legs and stocky shoulders) and his rear quarters look like the sleek leopard (long tail).

Lepjags have been kept at Waldo Animal retirement home near Gainesville, Florida and there are still some in captivity in the USA. Lepjags were owned by the magicians Siegfried and Roy. Leopard-jaguar hybrids are more massive than leopards. Their use as animal actors suggests that they are more manageable than purebred jaguars. Current captive lepjags are animal actors that have been retired. Although the term "lepjag" has currency in the USA, a German publication in 1978 refers to the male leopard x female jaguar hybrid as a leguar.

 

Siegfried and Roy had two spotted lepjags, Ali and Chico, and a black lepjag Sabaka, in their show in the 1990's, and listed them as lepjags in the show's 1994 program. However, in the 1998 souvenir program, they were listed as jaguars. In the book, "Siegfried and Roy - Mastering the Impossible" a photo on page 132 shows Siegfried and Roy in the pool with Ali who is identified on that page as being a lepjag. There is also a picture on page 260 of Siegfried and Roy with one of the lepjags in the show. They used them in the illusion where Roy (or rather a double) and a lepjag were shot out of a cannon; the second spotted lepjag was used to create the illusion. Kevin Chambers saw that show, and could tell right away that the animal that placed in the cannon was not the same as the one on the other side of the stage (where they had been shot to).

Mary Ann Howell and her husband visited the Secret Garden when Chico was on exhibit and were speaking with a security guard when another visitor asked why Chico had jaguar spots on his body, but leopard spots on his neck. The security guard looked at her list of animals, and confirmed that Chico was a lepjag hybrid.

In January 2010, the Serengetti zoological park in Sonora, Mexico, reported a new litter of cubs including a lepjag. They were described by the news as "one jaguar, one panther [leopard], and one jaguar-panther cross".

LIJAGULEP (CONGOLESE SPOTTED LION)

In 1908, a "Congolese Spotted Lion" was exhibited by a showman in London. It turned out to be the offspring of a female leopard/jaguar cross which was then crossed with a lion. It had been bred at Lincoln Park Zoo, Chicago, America (not Brookfield Zoo as this didn't open until 1934) and exported as a public attraction. The "manufacture" of supposedly exotic cats was once big business and exotic or freakish big cats were exhibited to the public with some story about them being a newly discovered animal. According to "Wild Cats Of The World" by Guggisberg, "Around the turn of the century three cubs were born at a Chicago zoo from a male jaguar and a female leopard. They were sold to a traveling menagerie and the male was killed by a lion, while the females grew to jaguar size. They were mated to a lion, which they seemed to prefer to a leopard, and produced several litters. One of these jaguar-leopard-lions came to London Zoo. The animal looked like a young, slim lioness but was marked with brown spots of a jaguar- or leopard-like pattern.". Guggisberg cites Pocock, R.I. 1908 "Hybrid Lion and Leopard" and The Field, April 18; Pocock, R.I. 1908 "The Supposed Lion and Leopard Hybrid." Others described it as being "the size of a lioness, but scrawny, with brown spots of either the jaguar or leopard variety."

Frohawk's drawing of the supposed lion-leopard at London Zoo, 1908, claimed to be a Congolese Spotted Lion and later revealed to be a lijagulep

The Times (April 15, 1908) pg. 6 stated "A Strange Animal From The Congo: Mr. J. D. Hamlyn, the animal dealer of St. George St.,E., who obtained two or three new monkeys from the Congo, has just recieved from the same region a very curious feline animal nearly as large as an adult lioness, which it resembles in build, but irregularly spotted. There is no trace of a mane or ruff, nor is the tail tufted as in the lion. The general hue is tawny, but with a rufous tinge, reminding one of the coat of a cheetah rather than of the leopard, and the inner sides of the limbs are yellowish white, with dark spots. The markings on the upper surface differ greatly in size and character; on the hind limbs they are large; toward the forequarters and head they diminish in size, but increase greatly in number, and the face is so to speak, strippled with black, except on the nose. There is a black mark on each side of the lower jaw, and a black stripe on the posterior side of each ear; and along the spine, from the root of the tail to about the centre of the back is a row of dark markings, somewhat like disconnected links of a chain. The hue of the tail for the greater part of it's length corresponds to that of the body, but the terminal portion is banded with black and white. The animal, a female, is in excellent condition and fairly quiet. The obvious suggestion is that the animal is a wild-bred hybrid, with a lioness for dam and a spotted cat for sire. Lion-tiger hybrids were bred in this country by Atkins, the proprietor of a famous travelling menagerie; among Continental breeders Carl Hagenbeck has been most successful. A cross between a puma and leopard has also been obtained, but wild-bred hybrids between the larger cats are exceedingly rare. Today the animal will be sent to the Zoological Gardens, where the question of its parentage will be scientifically investigated."

RI Pocock's letter to The Field (April 18th, 1908): "Sir - Since the days of Linnaeus and Schreber no large member of the cat tribe (Felis) alleged to have been caught or killed in the wild state has been imported into Europe which could not be at once assigned to one or other of the following six forms: lion, tiger, jaguar, leopard, ounce, or puma. Each of these represents a well-marked, easily recognisable type, and, despite their subdivision into numerous local races, every schoolboy can tell the main species apart without reference to geographical distribution. Of exceptional interest, therefore, is it to record that on Saturday last, the Zoological Society was informed that Mr Hamlyn, the well-known dealer in the east of London, had procured a large cat, exceeding a leopard in size, which he could not identify with any of the species just mentioned. Examination of the animal, which is now on exhibition in the lion house in the Zoological Gardens, fully justifies Mr Hamlyn's perplexity.

A superficial glance suggests that she is a hybrid between a lion and a leopard. She is larger than a leopard, and, although certainly adult, much smaller than a full-grown lioness. She stands about 30in at the shoulder, and is lank and long-limbed, approaching in build a lean lioness rather than a leopard, and resembling also the former rather than the latter in the shape of the head. The ground colour is tawny, becoming richer in tone in the posterior than in the anterior part of the body. On the throat and underside the tawny colour gradually fades into white. The whole animal is spotted like a leopard, but the spots are brown, not black, with the exception of some of the solid spots low down on the hind legs and on the belly. On the sacral region the spots are also nearly black, elongated, and arranged in longitudinal lines, and on the sides of the body they form rosettes as large as those of some jaguars, with slightly darker centres, containing smaller isolated spots. The tail is very remarkable. It is as long as is a leopard, and exceeds that of lion, the end being upcurved to keep clear of the ground. For the greater part of its length it is very short-haired and very faintly spotted, but the terminal 6in are longer haired, black-spotted, whitish beneath, and black, but not tufted at the tip. In short the distal end of the tail is like that of a leopard, and the proximal portion like that of a lion. Another leopard-like feature is the presence of a patch of black hair round the corner of the mouth, and of a transverse black stripe across the throat.

It appears to me that two, and only two, opinions can be held about this animal. She is either a lion-leopard hybrid or an example of a new species standing midway between these two species. I strongly incline to the former hypothesis, which would need no justification were the history of this animal known. The disparity in size between a large male leopard and a small lioness that had just reached maturity would not necessarily be an obstacle to pairing, and since it is known that lions and tigers on the one hand, and leopards and pumas on the other, have been successfully crossed in captivity, there is no reason for thinking that leopards and lions should not also be cross-bred; for, as I have recently pointed out, the relationship between leopards and lions is much closer than that between leopards and pumas, lions being in all probability directly descended from some species of Felis closely akin to leopards and jaguars, as is attested by the patter of the cubs, as well as by other points of resemblance. But the history of this animal cannot be squared with the supposition that she is a menagerie-bred hybrid, unless - which is hardly likely - she was bred by natives or traders in West Africa. According to the story told to Mr Hamlyn, she was brought as a cub by natives to a French trading settlement somewhere up-country from the Gaboon, and was kept in captivity for about two years before being transported to the west coast and shipped to Europe this spring in a French boat.

Now leopards, although apparently not common in the French Congo, seem to be almost ubiquitous in the forest region of West Africa. Lions, on the other hand, are unknown in the southern portions of that territory, although they have been recorded from its northern districts up towards Lake Chad. It is possible, therefore, that a hybrid between the two species may have been produced without human intervention in the northern area of French Congoland. This, however, seems highly improbably, for lions habitually frequent open, scrubby country, while leopards are essentially forest dwellers, and even if representatives of the two species were to meet, the encounter would more likely end in the death of the leopard that in the establishment of friendly relations between them. In addition to these objections must be urged the total absence up to the present time of records of the occurrence of natural hybrids, or of specimens suggesting in the remotest degree that they might be hybrids, between any two of the known wild species of Felis.

Against the hypothesis that the animal, which is certainly not a leopardess, and equally certainly not a lioness with retained cub spots, represents a large undescribed species of the cat tribe there is nothing to be urged beyond the circumstance that the existence of such a creature has never been hinted at or suspected, although good collections of the fauna of the country whence it is alleged to have come have reached the Paris and European museums. Possibly such a form may have been overlooked, but until the history of this specimen, as told to Mr Hamlyn by its original owner, has been verified it would be rash to regard it as an example of a genuine wild species. All that can be done at present is to urge naturalists and sportsmen who visit the French Congo in the future to keep a sharp lookout for a second specimen of this most interesting beast."

In The Field No 2887, April 25th, 1908, the editor wrote "An illustration reproduced from a drawing by Mr F W Frohawk, of the supposed lion-leopard at the Zoological Gardens is now presented, and it will be interesting to compare it with the picture which accompanies the letter on feline hybrids by Mr Scherren. Mr R I Pocock's remarks on this interesting animal which appeared in last week's "Field" leave little more to be said at present. It would certainly appear to be either a hybrid lion-leopard, or else a new species of large leopard, a supposition strengthened by several points of closer resemblance to a leopard that to a lion, and the pattern of the larger rosette markings which are like those of the snow leopard (Felis uncia). It may be well to note in further detail the colouring and markings of the beast. The ground colour is a pale tawny-buff, blending into creamy-white on the undersurface of the body; chin, throat, chest, inside of legs and undersurface of end of tail whitel; the whole surface of the body and legs is spotted similar or a leopard and snow leopard, the head and neck being less plainly marked; all the markings on the upper parts are pale, dusky, developing into black below, and deep black on both surfaces of the legs. An important feature is the pattern of the larger rosette markings, which are similar to those of the snow leopard, being composed of smaller spots, but forming larger rosettes than is usual in the ordinary leopard. Excepting the crown, the spots on the head, and especially on the neck, are small, and more or less indistinct. The fore paws, rump and basal three-fourths of the tailare much like those of a lioness in form, but the end of the tail, although less ample, is marked like a show leopard's. The black angle of the nmouth is similar to that of a leopard; the nose is dull-pink and eyes pale ochreous, like most leopards; but the general squareness of the head and rather large ears are more lion like. At times, when the animal is standing slackly, she is hollow backed, but usually the back is as shown in the drawing. That idea that this animal may possibly prove to be a new species of great cat will not be generally entertained; but it must be remembered that a far more conspicuous creature, to wit, the okapi, has only been made known to us of late years, and that it is possible that such an animal as that now in question frequenting, as it would undoubtedly do, dense forest regions, and being of nocturnal habits, might have escaped observation."

The possibility was considered that the Congolese spotted lion may have been part cheetah or a new species of leopard. In The Field No 2887, April 25th, 1908, Henry Scherren wrote "In all probability the interesting animal now in the lion house of the Zoological Gardens is the only feline hybrid yet exhibited for which the claim has been advanced that it was wild bred. The story of its origin as told to Mr Hamlyn, and given by Mr Pocock in his letter,is of considerable interest; but, in my opinion, the interest will be heightened when the gentleman by whom the animal was consigned to Mr Hamlyn gives us full particulars. On the view that this animal was bred in the Congo, with a lioness for dam, there could have been but two possible sires: the leopard or the cheetah. Mr Pocock has given his reasons for accepting the former, but I think he will admit that there is a superficial resemblance to the latter. Conclusive evidence against the supposition that a cheetah had any share in the parentage - tough it occurred to me when I first saw the animal in her travelling box - is afforded by the size of the head, the massive forelimbs, and the retractile claws.

Certain difficulties, however, present themselves with regard to the story that the animal was wild bred. One was pointed out by Mr Pocock in hias remark that 'if representatives of the two species were to meet the encounter would be more likely to end in the death of the leopard than in the establishment of friendly relations between them.' next I cannot equate the apearance of the animal in regard to age and development with the scanty details that have been given to Mr Hamlyn. Granting that it was even two years old when brought by the natives to the French trading settlement, the two years spent in confinement in Africa and the time occupied in the passage to Europe do not, in my opinion, account for the whole span of its existence. And from what I saw of the hybrid before she was unpacked, and afterwards in one of the spacious dens in the lion house, I should have come to the conclusion that she was well used to being exhibited but for the assurance from Mr Hamlyn that this was not the case. Everybody who has seen the animal will agree with Mr Pocock that it is of the highest interest, and it is to be hoped that it will remain in its present quarters." The markings on the Congolese spotted lion in Frohawk's illustration (The Field No 2887, April 25th, 1908) exactly match those on the museum specimen.

An article from The Times for Monday May 4, 1908 pg. 12 said "Sale Of A Supposed Congo Hybrid. After having been on view in the lion house at the Zoological Garden for about a fortnight the feline hybrid described in The Times of April 15 was sold by auction at Aldridge's on Saturday. The attendance was very large; among those present were a good many showmen. Bidding began at 100 guineas, and eventually the animal was knocked down by Mr. Bostock at 1,030 guineas. One of the conditions of sale appears somewhat strange for it disclaimed any guarantee as to the animal's breeding, age, or any other description. This would seem to show that the story of the animal, as told to Mr. Hamlyn by the original owner, has not been verified. The story was that the hybrid had been brought as a cub by natives to a French trading settlement somewhere up country from the Gaboon, and kept in captivity for about two years before being transported to the West Coast and shipped to Europe in a french boat. At any rate the responsible officials at the Zoological Society do not appear to have been convinced by the story, or they would probably have made some offer for the animal, for which about 500 pounds was asked a fortnight ago. The keepers in the lion house maintain a strong opinion that it was bred in a menagerie; the responsible officials are more reticent on the subject and prefer to wait for evidence as to the place of birth, and the species from which the creature was bred."

In The Field No 2889, May 9th, 1908, R I Pocock wrote of "The Supposed Lion-Leopard Hybrid" "Sir - Since you were good enough to publish in the Field of April 18 my description of the supposed lion-leopard hybrid with its history as issued to the press by Mr Hamlyn, I should like to give what I am convinced is the true sotry of its origin and antecedents, so as to lay at rest once and for all the idea that it was a natural product of the French Congo.

Some years ago, three hybrids, one male, two female, were bred in Chicago from a male jaguar and an Indian leopardess, and were bought by the proprietor of an American traveeling show of performing animals. The male was killed by a lion, but the females lived, grw to the size of a jaguar, and when adult were mated with a young lion, choosing him, it is said, in preference to male leopards. Several litters were born, each consisting of two cubs. These resembled a lion in general colour, but were spotted. In the case of every litter the spots of one cub were like those of the jaguar, and the otherlike those of a leopard. The males were without mane. At the end of last year, some of these animals, then about three and a half years old, were alive in the United States.

These facts I can vouch for on first hand authority. My conviction that the animal recently exhibited in the Zoological Gardens is one of those hybrids with jaguar-like spots is a conclusion deduced from a combination of circumstances, partly from a knowledge of the recent importation by Mr Bostock from America of a number of animals for the exhibition at Earl's Court, partly from a clue supplied to me by Mr Carl Hagenbeck, who predicted almost to the letter the outcome of the sale, partly from overheard remarks let drop at the auction at Aldridge's, and finally from the fact that the animal was knocked down to Mr Bostock for a sum representing ten times its market value. With the above-mentioned facts before them, your readers will be able to piece in the details of the entire transaction without further comment on my part. From a scientific standpoint the animal gains interest from a knowledge of its true nature. Not suspecting three species to be involved in its parentage, I was not quite right in determining it as a lion-leopard hybrid, although the spots, as I stated, obviously recall those of a jaguar, I dismissed that species in considering its pedigree on account of its comparatively slender build and the great length of the tail. He elimination of the shortness of the tail and of the sturdiness in shape of the jaguar is not surprising, however, seeing that these characters are only found in one out of the three parent forms. A further point of interest to be noted in this. The extreme closeness of the affinity between leopards and jaguars, which has already been claimed on other grounds, is further attested by their hybrids forming fertile unions with a species quite distinct from either of the parents. I learn from Mr hagenbeck that lion-tiger hybrids appear to be sterile, not only when paired with one another, but also when mated with the parent species. At all events one of his female hybrids produced no cubs after being mated with a male lion." (In this Pocock was incorrect as it was subsequently found that female ligers and tigons can produce young when mated to a male tiger or lion.)

The fate of the "Congolese Spotted Lion" (the lijagulep exhibited in Britain) is mentioned in R I Pocock's letter to The Field of 2nd November 1912 where he compared it to the leopons born in Kolhapur (1910): "The nearest approach to [a lion-leopard] hybrid hitherto reported is the one bred at Chicago between a male lion and a female cross between a jaguar and a leopard, the true story of which, accompanied by a good figure by Mr Frohawk, may be found in the Field for April 18 and 25, and May 9, 1908. The final episode in the history of that animal has, I believe, not yet been told. After being exhibited in the Zoological Gardens and at the White City it went to Glasgow, where, according to a sensational Press notice, it was killed by a lion, which broke down the partition between the cages and made short work of its opponent. That this story was of a piece with the original account of the hybrid given out when it first appeared on the market may be inferred from the condition of the dressed skin, which had no sign of a tear or scratch upon it in London shortly after the alleged tragedy. The chief difference between this hybrid of three species and the lion-leopard born at Kolhapur lies in the size of the spots, those of the [lijagulep] being large and jaguar-like, as might be expected, while those of the [leopon] are small and more leopard-like."

From Pocock's comments, the hybrids appear to have been born around 1904. There were probably at least 3 litter born with the lion siring cubs on both female lep-jags. SS Flower (1929) described London Zoo's hybrid (exhibited 1908) between a male lion and female leopard/jaguar hybrid.

BLACK JAGLIONS

Black jaguars (black panthers) have always caught the public imagination. The gene for melanism in jaguars is incompletely dominant hence "black" jaglions, having only one copy of the gene, are charcoal grey rather than glossy black.

INTRA-SPECIFIC JAGUAR HYBRIDS

Panthera onca onca x P o palustris – 3 male hybrids born Brasilia, Brazil in 1966. International Zoo Yearbook (1968)

REFERENCES AND FURTHER READING

Historical accounts (chronological order): R De Davison (1863), Sacc (1863), A Rörig (1903 – quotes Sacc), H Scherren (1908), SS Flower (1929), O Antonius (1951), H Petzsch (1956), H Hemmer (1966, 1968), H Windischbauer (1968), C Pohle (1969).

Helmut Hemmer: "Report on a Hybrid Between Lion x Jaguar x Leopard - Panthera leo x Panthera onca x Panthera pardus" in Saeugetierkundliche-Mitteilungen, 1968; 16(2): 179-182.
Howell, Mary-Anne: Personal Correspondence 2006/7
G Peters included several hybrids (liger, tigon, leopon, leguar) in his "Comparative Investigation of Vocalisation in Several Felids" published in German in Spixiana-Supplement, 1978; (1): 1-206.
Pocock, R.I. 1908 "Hybrid Lion and Leopard
The Times for Monday May 4, 1908 pg. 12
The Times (April 15, 1908) pg. 6

Textual content is licensed under the GFDL.

Many thanks to Paul McCarthy for tirelessly researching back issues of The Field and The Times to unravel the case of the Congolese Spotted Lion. .

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